Faction:
All
Levels:
11 to 15 for Federation
30 to 32 for Klingon and Romulan
Missions:
1) "Skirmish"
2) "Spin the Wheel"
3) "What Lies Beneath"
4) "Everything Old is New"
5) "Night of the Comet"
Federation:
Federation players get this story arc early as it merges with another story arc of which the Federation players receive that other factions do not which rewards this achievement:
Klingon and Romulan:
Simply play through your story missions until you get given this story arc.